四川动物  2015, Vol. 34(5) 688-694

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方福康, 王军旺, 陈元晓
FANG Fukang, WANG Junwang, CHEN Yuanxiao
中国喇叭螺属的研究及一新种记述
Studies on the Genus Boysidia Ancey from China, with Description of One New Species (Pulmonata: Stylommatophora: Pupillidae)
四川动物, 2015, 34(5): 688-694
Sichuan Journal of Zoology, 2015, 34(5): 688-694
10.11984/j.issn.1000-7083.20140580

文章历史

收稿日期:2014-12-24
接受日期:2015-04-08
中国喇叭螺属的研究及一新种记述
方福康1, 王军旺2, 陈元晓1     
1. 昆明医科大学细胞生物学与医学遗传学系, 昆明 650500;
2. 浙江清凉峰国家级自然保护区管理局, 浙江临安 311321
摘要:笔者整理和厘清了Boysidia Ancey,1881已记载的中国喇叭螺属的种类,共计24种,其中有1新种,即清凉峰贝喇叭螺新种Boysidia(Bensonella) qingliangfengensis sp. nov.,对喇叭螺各亚属、贝喇叭螺亚属的已知种列出检索表,并对新种的形态特征、栖息环境进行了记述,与相似种进行了讨论。新种模式标本分别保存在中国科学院动物研究所标本馆和昆明医科大学细胞生物学与医学遗传学系标本室。
关键词腹足纲     肺螺亚纲     柄眼目     虹蛹螺科     喇叭螺属     新种    
Studies on the Genus Boysidia Ancey from China, with Description of One New Species (Pulmonata: Stylommatophora: Pupillidae)
FANG Fukang1, WANG Junwang2, CHEN Yuanxiao1     
1. Department of Cell Biology and Medical Genetics, Kunming Medical University, Kunming 650500, China;
2. Management Bureau of Zhejiang Qingliangfeng National Nature Reserve, Lin'an, Zhejiang Province 311321, China
Abstract:Boysidia Ancey, 1881 is a small group in Chondrininae and distributes in South and Southeast Asia. A total of 24 species were recorded in south Qinling Mountains, China. Among of these species, one was identified as a new record. Descriptions of this new species and other 8 species of Boysidia (Bensonenella) were given in the text. The type specimens of the new species were kept in Institute of Zoology, Chinese Academy of Sciences, and the Department of Cell Biology and Medical Genetics, Kunming Medical University.
Key to subgenus of Boysidia (Boysidia), Boysidia (Bensonella), Boysidia (Paraboysidia) and Boysidia (Dasypupa)
1. Columellar tooth with node-shaped on aperture deep, angular tooth and parietal tooth united into 2 crescent knife-shaped tooch......Boysidia (Dasypupa) Thompson and Dance, 1983
Columellar tooth without node-shaped on aperture deep......2
2. Columellar tooth oblique......3
Columellar tooth not oblique, horizontal......Boysidia (Paraboysidia) Pilsbry, 1917
3. Angular tooth and parietal tooth united into 1 flake-shaped tooch, palatal tooth with hook-shaped......Boysidia (Bensonella) Pilsbry, 1900
Angular tooth and parietal tooth independence, parallel......Boysidia(Boysidia)Pilsbry, 1917
Boysidia (Bensonella) qingliangfengensis sp. nov. (Fig. 1: A-E)
Holotype: alt. 3.40 mm, diam. 2.10 mm; alt. of aperture 1.15 mm, diam. of aperture 0.75 mm. Collected from Longtangshan town, Linan city, Qingliangfeng Nature Reserve (30°01'N, 118°80'E), June 26, 2013.
Paratypes: 126 specimens. alt. 3.30-3.45 mm, diam. 2.00-2.20 mm; alt. of aperture 1.10-1.20 mm, diam. of aperture 0.700-0.815 mm. Collected from Longtangshan town, Linan city, Qingliangfeng Nature Reserve (30°01'N, 118°80'E), June 26, 2013.
Etymology: The name of the new species was based on Longtangshan town, Linan city, Qingliangfeng Nature Reserve from where the specimens were collected.
Shell small, thin, luster, elongatedly conic. 5(1/2) whorls, moderately convex, spire long, the body whorl inflated, the bottom of body whorl not extended to aperture, and not form a bell-shaped. Apex blunt, embryo-whorls (1-2(1/2)) increased slowly, smooth, striae absent. Suture deep. Shell yellow brown or pale yellow in colour. Aperture elliptical, aperture margin thick, slightly expanded and not reflexed, with 16 rib-strias on the neck of aperture. With 14 teeth in the aperture: 1 little nipple-formed infraparietal lamella, l large flake-formed parietal lamalla, 1 big flake-formed angular lamella, 1 small nipple-formed twin-lamella, 2 small nipple-shaped parallel lamella on right aperture margin, 2 small nipple-shaped suprapalaral plicae on the parallel lamella down, 1 big nipple-shaped upper palatal plicae and 1 small nipple-formed interpalatal plicae, 1 small nipple-shaped lower palatal plicae on aperture botton. 1 big nipple-formed basal plicae on the below part of columella margin nearly, 1 small nipple-shaped columellar lamella on the columella margin above, 1 small nipple-formed subcolumellar lamella on the columellar lamella down. Umbilicus big, with hole-formed, and the umbilicus was not covered by columella margin.
Habitat: Boysidia (Bensonella) qingliangfengensis sp. nov., as other Boysidia (Bensonella) species, usually lives on limestone areas of mountainous regions, in dark, moist habitats with bryophyte and lichen, in the cracks of the rocks or under the faller leaves.
This new species is bigger than Boysidia (Bensonella) plicidens (Benson, 1849), but differs obviously from the bigger size: the new species shell with whorls 5(1/2), alt. 3.40 mm, diam. 2.10 mm; alt. of aperture 1.15 mm, diam. of aperture 0.75 mm. Aperture elliptical, aperture margin thick, slightly expanded and not reflexed, with 16 rib-strias on the neck of aperture, with 14 teeth on the aperture: 1 little nipple-formed infraparietal lamella, l large flake-formed parietal lamalla, 1 big flake-formed angular lamella, 1 small nipple-formed twin-lamella, 2 small nipple-shaped parallel lamella on right aperture margin, 2 small nipple-shaped suprapalaral plicae on the parallel lamella down, 1 big nipple-shaped upper palatal plicae and 1 small nipple-formed interpalatal plicae, 1 small nipple-shaped lower palatal plicae on aperture botton. 1 big nipple-formed basal plicae on the below part of columella margin, 1 small nipple-shaped columellar lamella on the columella margin above, 1 small nipple-formed subcolumellar lamella on the columellar lamella down. The latter with 5 whorls (alt. 2.00 mm, diam. 1.50) in small size, and the aperture irregular, subtriangular, having 9 folds teeth: 3 parietal lamellae, of which the upper two are long; one small dentiform columellar lamellae; five palatals, of which two subbasal plicae being minute. Basal margin has a callous outside. Umbilicus narrow.
Key words: Pulmonata     Stylommatophora     Pupillidae     Boysidia (Bensonella)     new species    

虹蛹螺科Pupillidae喇叭螺属Boysidia由Ancey于1881年创立,其主要特征为:贝壳较小,呈圆锥形,具有狭窄的脐孔,一般具有5~6 个膨大的螺层,体螺层的笔直延伸,有时向下弯曲;在大多数情况下壳口内具有6枚大齿,壳口稍垂直,有时有附属的小齿存在,或基褶缺乏,独立的角板和壁板,或愈合成一弯曲的板。口缘十分扩大。

本属种类主要分布于我国黄河流域以南的广大地区和南亚、东南亚各国。1931年Thiele将此属分为2个亚属,即Boysidia (Boysidia)喇叭螺属指名亚属和Boysidia (Bensonella)贝喇叭螺亚属。指名亚属的特征是在倒数第二螺层壁板和角板融合成1个叶板,轴板在内向下弯曲。贝喇叭螺亚属的特征是壁板和角板分离,柱板水平状,不在内向下弯曲。Pilsbry(1916—1918)将Boysidia分为3个亚属,即:Boysidia (Boysidia)喇叭螺属指名亚属、Boysidia (Bensonella)贝喇叭螺亚属和Boysidia (Paraboysidia)Pilsbry,1917奇喇叭螺亚属。奇喇叭螺亚属的种类壳口上的齿不呈钩状,如Boysidia (Paraboysidia)paviei Bavay & Dautzenberg,贝喇叭螺亚属壳口上的齿外表呈钩状,如Boysidia (Bensonella)plicidens (Benson)。Thompson和Dance于1983年又创建一新亚属,即毛蛹喇叭螺亚属Boysidia (Dasypupa),此亚属仅见于东南亚国家。

喇叭螺属Boysidia Ancey,1881

Boysidia Ancey,1881 Le Naturalisle,p. 373.

Gredeileriella Moellendorff,1884

Dasypupa Thompson and Dance, 1983

Paraboysidia Pilsbry,1917

Boysidia Pilsbry,1916-1918 Manual of Conch.,24:192-209.

Boysidia Thiele,1931 H and ,System.,p. 830-831.

Boysidia Yen,1939 Abh. Mus. Nat. Ges.,444: 75.

Boysidia Chen and Gao 1987 Sci. Press Beijing. China.,p. 43-44.

Boysidia Abbott and Boss, 1989

Boysidia Panha and Burch, 2004—2005

模式种:Pupa dorsata Ancey,1881 Le Naturalisle,p.373. 喇叭螺属指名亚属Boysidia(Boysidia)Pilsbry,1917 贝喇叭螺亚属Boysidia(Bensonella) Pilsbry,1900 奇喇叭螺亚属Boysidia(Paraboysidia)Pilsbry,1917 毛蛹喇叭螺亚属Boysidia(Dasypupa)Thompson and Dance, 1983

喇叭螺属各亚属检索表

1. 轴板呈结节状,深入壳口中;角板和壁板融合为一短的,弯曲的两叶刀形 毛蛹喇叭螺亚属Boysidia(Dasypupa)Thompson and Dance, 1983 轴板不呈结节状2 2. 柱板向下倾斜3 柱板进入部分向内呈水平状,不向下倾斜奇喇叭螺亚属Boysidia(Paraboysidia)Pilsbry,1917 3. 角板和壁板融合为一两叶板或单独板,壳口内腭褶和板呈钩状 贝喇叭螺亚属Boysidia(Bensonella) Pilsbry,1900 角板和壁板独立、平行喇叭螺亚属Boysidia(Boysidia)Pilsbry,1917

喇叭螺属指名亚属Subgenus Boysidia(Boysidia)Pilsbry,1917

Boysidia(Boysidia)Pilsbry,1917 Man. Conch.,2(24): 194. 模式种:Pupa dorsata Ancey,1881 1. 囊喇叭螺Boysidia(Boysidia) dorsata(Ancey,1881)

Pupa dorsata Ancey,1881 Le Naturaliste.,p. 373.

Boysidia(Boysidia)dorsata Pilsbry,1917 Man. Conch.,2(24): 195-196.

Boysidia dorsata Yen,1939 Ahb. Sencken. Nat. Mus. Ges. 444: 75,pl. 6. f. 39.

Boysidia(Boysidia)dorsata Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国江西省鄱阳湖。

分布:江西省。 2. 湖南喇叭螺Boysidia(Boysidia)hunana(Gredler,1881)

Pupa hunana Gredler,1881 Jahrb. D. Deutschen. Malak. Ges.,8: 23,pl. 1. f. 5.

Pupa hunana Heude,1882 Mem. D’Hist. nat. l’Emp. Chinois.,p.76,pl. 18. f. 25.

Pupa(Guredleriella)hunanensis Gredler,1884 Jahrb. D. Deutschen. Malak. Ges.,6: 179.

Boysidia(Boysidia)hunana Pilsbry,1916-1918. Man. Conch.,2(24): 194,pl.33,f. 8,9.

Boysidia hunana hunana Yen,1939 Ahb. Sencken. Nat. Mus. Ges. 444: 76,pl. 6. f. 40.

Boysidia(Boysidia)dorsata Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国湖南省岳阳市。

分布:湖南省西北部、湖北省西北部及长江流域。 3. 显著喇叭螺Boysidia(Boysidia) conspicua(Moellendorff,1885)

Pupa(Boysidia) hunanensis var. conspicua Moellendorff,1885 Jahrb. D. Deutschen. Malak. Ges.,7: 396.

Boysidia(Boysidia) hunanensis conspicua Pilsbry,1916-1918. Man. Conch.,2(24): 194-195,pl.33,f. 10-12.

Boysidia hunana conspicua Yen,1939 Ahb. Sencken. Nat. Mus. Ges.,444: 76,pl. 6. f. 41.

Boysidia(Boysidia) conspicua Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国广东省肇庆市。

分布:广东省。 4. 弯曲喇叭螺Boysidia(Boysidia)strophostoma(Moellendorff,1885)

Pupa(Boysidia)strophostoma Moellendorff,1885 Jahrb. D. Deutschen. Malak. Ges.,7: 395,pl. 11,f. 23.

Boysidia(Boysidia)strophostoma. Pilsbry,1916-1918. Man. Conch.,2(24): 197-198,pl. 33,f. 15,16.

Boysidia strophostoma Yen,1939 Ahb. Sencken. Nat. Mus. Ges.,444: 76,pl. 6. f. 42.

Boysidia(Boysidia) conspicua Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国广东省肇庆市。

分布:广东省、广西省、湖南省、湖北省。 5. 细纹喇叭螺Boysidia(Boysidia)gracilisHaas,1937

Boysidia(Boysidia)gracilisHaas,1937 Arch. Moll.,69: 7,f. 18-20.

Boysidia gracilisYen,1939 Ahb. Sencken. Nat. Mus. Ges.,444: 76,pl. 6. f. 43.

Boysidia(Boysidia)gracilis,Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国湖北省巴东县。

分布:陕西省、四川省、湖北省、重庆省。 6. 杭州喇叭螺Boysidia(Boysidia) hangchowensis(Pilsbry & Hirase,1908)

Boysidia(Boysidia)hangchowensis Pilsbry & Hirase,1908Proc.Acad. Nat. Sci.,p. 42,f. 6.

Boysidia(Boysidia)hangchowensis Pilsbry,1916-1918 Man. Conch.,24: 196-197,pl. 33. f. 17.

Boysidia(Boysidia)hangchowensis Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国浙江省杭州市。

分布:浙江省。 7. 凤县喇叭螺Boysidia(Boysidia)fengxianensis Chen & Wu,1999

Boysidia(Boysidia)fengxianensis Chen & Wu,1999 Acta Zootax. Sinica.,24(2): 127-135. f. 1.

Boysidia(Boysidia)fengxianensis Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国陕西省凤县。

分布:陕西省。 8. 五齿喇叭螺Boysidia(Boysidia)pentadensChen & Wu,1999

Boysidia(Boysidia)pentadens, Chen & Wu,1999 Acta Zootax. Sinica.,24(2): 127-135. f. 2.

Boysidia(Boysidia)pentadens, Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国云南省勐腊县勐仑。

分布:云南省。 9. 西山喇叭螺Boysidia(Boysidia)xishanensis Chen & Wu,1999

Boysidia(Boysidia)xishanensis Chen & Wu,1999 Acta Zootax. Sinica.,24(2): 128-135. f. 3.

Boysidia(Boysidia)xishanensis Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国云南省昆明市西山。

分布:云南省。 10. 太白喇叭螺Boysidia(Boysidia)taibainensis Chen & Wu,1999

Boysidia(Boysidia)taibainensis Chen & Wu,1999 Acta Zootax. Sinica.,24(2): 128-135. f. 4.

Boysidia(Boysidia)taibainensis Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国陕西省太白县。

分布:陕西省。 11. 黄果树喇叭螺Boysidia(Boysidia)huangguoshuensis Luo & Chen,2000

Boysidia(Boysidia)huangguoshuensis Luo & Chen,2000 Acta Zootax. Sinica.,25(2): 148-151. f. 5-7.

Boysidia(Boysidia)huangguoshuensis Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国贵州省镇宁布衣族苗族自治县黄果树。

分布:贵州省。 12. 咸丰喇叭螺Boysidia(Boysidia)xianfengensis Zhang & Chen,2014

Boysidia(Boysidia)xianfengensis Zhang & Chen,2014 Acta Zootax. Sinica.,39(4): 570-575. f. 1-5.

模式标本产地:中国湖北省咸丰县城关。

分布:湖北省。 13. 小关喇叭螺Boysidia(Boysidia)xiaoguanensis Zhang & Chen,2014

Boysidia(Boysidia)xianfengensis Zhang & Chen,2014 Acta Zootax. Sinica.,39(4): 570-575. f. 6-10.

模式标本产地:中国湖北省宣恩县小关。

分布:湖北省宣恩县、恩施县。 14. 巩尧山喇叭螺Boysidia(Boysidia)gongyaoshanensis Yang & Zhang,2012

Boysidia(Boysidia)gongyaoshanensis Yang & Zhang,2012 Acta Zootax. Sinica.,37(3): 550-551. f. 1-3.

模式标本产地:中国贵州省兴义市巩尧山。

分布:贵州省。

喇叭螺属贝喇叭螺亚属Subgenus Boysidia (Bensonella) Pilsbry,1900

1. 皱褶贝喇叭螺Boysidia (Bensonella) plicidens(Benson,1849)

Pupa plicidens Benson,1849 Ann. Mag Nat. Hist.,2(4): 126.

Boysidia (Bensonella) plicidens Pilsbry,1916-1918 Man. Conch.,2(24): 198-199,pl.34,f. 1-4,9,10.

模式标本产地:印度。

分布:中国浙江省、印度、日本。 2. 双片贝喇叭螺Boysidia (Bensonella)dilamellaris Chen & Wu,1999

Boysidia (Bensonella)dilamellaris Chen & Wu,1999 Acta Zootax. Sinica.,24(2): 130-135. f. 5.

Boysidia (Bensonella)dilamellaris Chen and Zhang, 2002 Acta Zootax.,27(4): 693.

模式标本产地:中国陕西省安康市吉河。

分布:陕西省。 3. 石林贝喇叭螺Boysidia (Bensonella)shilinensis Chen & Wu,1999

Boysidia (Bensonella)shilinensis Chen & Wu,1999 Acta Zootax. Sinica.,24(2): 131-35. f. 6.

Boysidia (Bensonella)shilinensis Chen and Zhang, 2002 Acta Zootax.,27(4): 693.

模式标本产地:中国云南县石林县石林。

分布:云南省。 4. 天星桥贝喇叭螺Boysidia (Bensonella)tianxingqiaoensis Luo and Chen, 2000

Boysidia (Bensonella)tianxingqiaoensis Luo and Chen, 2000 Acta Zootax. Sinica.,252: 147-151,f. 1-4.

Boysidia (Bensonella)tianxingqiaoensis Chen and Zhang, 2002 Acta Zootax.,27(4): 693.

模式标本产地:中国贵州省镇宁布衣族苗族自治县天星桥。

分布:贵州省。 5. 潼关贝喇叭螺Boysidia (Bensonella)tongguanensis Chen & Zhang,2002

Boysidia (Bensonella) tongguanensis Chen & Zhang,2002 Acta Zootax.,27(4): 693-698,f. 1-4.

模式标本产地:中国陕西省潼关县李家乡。

分布:陕西省。 6. 修文贝喇叭螺 Boysidia (Bensonella)xiuwenensis Zhang &Luo,2010

Boysidia (Bensonella)xiuwenensis Zhang &Luo,2010 Acta Zootax.,35(4): 865-867,f. 1-3.

模式标本产地:中国贵州省修文县回水石林。

分布:贵州省。 7. 兴义贝喇叭螺Boysidia (Bensonella)xingyiensis Guo & Zhou,2006

Boysidia (Bensonella)xingyiensis Guo & Zhou,2006 Acta Zootax.,31(3): 541-543,f. 1-4.

模式标本产地:中国贵州省兴义县纳灰。

分布:贵州省。 8. 南江贝喇叭螺Boysidia (Bensonella) nanjiangensis Zhang & Zhang,2011

Boysidia (Bensonella) nanjiangensis Zhang & Zhang,2011 Acta Zootax.,36(3): 718-721,f. 1-3.

模式标本产地:中国贵州省开阳县南江峡谷。

分布:贵州省。

贝喇叭亚属分种检索表

1. 柱板进入部分向内呈水平状,不向下倾斜2 柱板向内下倾斜4 2. 壳高大于2 mm,螺层数612个 Boysidia (Bensonella)shilinensis Chen & Wu,1999 壳高小于2 mm,螺层数不超过512个3 3. 壳口具腭褶9枚 Boysidia (Bensonella) plicidens(Benson,1849) 壳口具腭褶2枚Boysidia (Bensonella)dilamellaris Chen & Wu,1999 壳口具腭褶5枚,有下腭褶 Boysidia (Bensonella)tianxingqiaoensis Luo and Chen, 2000 壳口具腭褶5枚,无下腭褶 Boysidia (Bensonella)tongguanensis Chen & Zhang,2002 4. 壳高小于3.3 mm5 5. 壳口完全分离,呈喇叭状Boysidia (Bensonella)xingyiensis Guo & Zhou,2006 壳口不完全分离,不呈喇叭状Boysidia (Bensonella)xiuwenensis Zhang &Luo,2010 6. 壳口内有6枚板与皱褶 Boysidia (Bensonella)nanjiangensis Zhang & Zhang,2011 壳口内有14枚齿,柱板和亚柱板不融合清凉峰贝喇叭螺 新种 Boysidia (Bensonella)qingliangfengensis sp. nov.

清凉峰贝喇叭螺新种 Boysidia (Bensonella)qingliangfengensis sp. nov.

正模标本

壳高3.40 mm,壳宽2.10 mm,壳口高 1.15 mm,壳口宽0.75 mm,2013年6月26日采自浙江省临安市清凉峰自然保护区龙塘山。

副模标本

126个,壳高3.30~3.45 mm。壳宽2.00~2.20 mm,壳口高1.10~1.20 mm,壳口宽0.700~0.815 mm。 2013年6月26日采自浙江省临安市清凉峰自然保护区龙塘山(30°01′N,118°80′E)。

词源

新种的种名源于本种的采集地。

形态特征

贝壳微小,壳质薄,有光泽,贝壳呈高圆锥形,螺旋部较高,体螺层膨大,其底部不向壳口延伸,不形成喇叭状,有5个螺层。壳顶钝,胚螺层(1~2个螺层)光滑,无螺纹。缝合线深。壳面呈黄褐色或淡黄褐色。壳口呈椭圆形,口缘稍厚、不外折、扩大,在壳口颈部具有16条肋纹,壳口内有14枚齿:即有1枚细小的呈乳头状的下壁板(infraparietal lamella),1枚较大呈片状的壁板齿(parietal lamalla),1枚较大呈片状的角板齿(angular lamella),有1枚较小呈乳头状的孪板(twin lamella),靠近右侧上口缘处有2枚较小,呈乳头状的平行板(parallel lamella),在壳口右侧口缘处有2枚较小,呈乳头状的上上腭褶齿(suprapalaral plicae),其下方有1枚较大的呈乳头状上腭褶齿(upper palatal plicae),其内侧有1枚较小呈乳头状的中腭褶齿(interpalatal plicae),在壳口底部内有1枚较小的呈乳头状的下腭褶齿(lower palatal plicae)。靠近轴缘下方,有1枚较大的呈乳头状的基褶齿(basal fold),其左上方有1枚稍小的呈乳头状的柱板齿(columellar lamella),其下方有1枚小的呈乳头状的亚柱板齿(subcolumellar lamella)。脐孔不被轴缘遮盖,脐孔大而深。
图 1 清凉峰贝喇叭螺新种 Fig. 1 Boysidia (Bensonella)qingliangfengensis sp. nov. A. 壳口放大aperture enlarge,B. 贝壳右侧面观right lateral view of shell,C. 贝壳侧面观lateral view of shell,D. 壳背面观dorsal view of shell,E. 贝壳腹面观ventral lateral view of shell.

栖息环境

生活在阴暗潮湿、多石灰岩地区,常栖息于潮湿、多地衣、苔藓的石灰岩石壁上,岩石缝隙中,树叶下。

讨论

新种与皱褶贝喇叭螺Boysidia(Bensonella)plicidens(Benson,1849)在外形上较相似,前者有5个螺层,壳高3.40 mm,壳宽2.10 mm,壳口高1.15 mm,壳口宽0.75 mm。壳口呈椭圆形,口缘稍厚、不外折、扩大,在壳口颈部具有16条肋纹,壳口内有14枚齿:即有1枚较大呈片状的壁板齿,1枚稍大较小呈片状的角板齿,有1枚较小,呈乳头状的上腭褶齿,靠近右侧口缘处有1枚较大的呈乳头状的上腭褶齿,在壳口底部有1枚较小呈乳头状的下腭褶齿,靠近轴缘下方,有1枚稍大呈乳头状的亚柱板齿,其上方有1枚较大的呈乳头状的柱板齿,1枚较小呈乳头状的上柱板齿和1枚稍大呈乳头状的下壁板齿。但后者较小(壳高2.00 mm,壳宽1.50 mm),有5个螺层,壳口不规则,近三角形,壳口内有9枚齿:有3枚壁板齿,其中上面2枚较长;1枚小齿状的柱板齿;5枚腭褶齿,其中2枚亚基褶齿较小。与前者有所区别。

喇叭螺属奇喇叭螺亚属Subgenus Boysidia (Paraboysidia)Pilsbry,1917

湖北奇喇叭螺 Boysidia (Paraboysidia)hupeina (Gredler,1901)

Hypselostoma hupeinum Gredler,1901 Nachr. Bl.,p. 151.

Boysidia hupeina Yen,1939 Ahb. Sencken. Nat. Mus. Ges.,444: 76,pl. 6. f. 44.

Boysidia (Paraboysidia)hupeina Chen and Zhang, 2002 Acta Zootax.,27(4): 692.

模式标本产地:中国湖北省北部。

分布:湖北西北部。

现已知我国虹蛹螺科喇叭螺属共计 24种,其中喇叭螺属指名亚属Boysidia (Boysidia)有14种,贝喇叭螺亚属Boysidia (Bensonella)有9种,奇喇叭螺亚属Boysidia (Paraboysidia)有1种,主要分布于我国浙江、广西、湖南、湖北、贵州、云南、四川、陕西和甘肃等省。长江流域和秦岭以南地区的种类尤喜栖息于石灰岩地区。但笔者发现此属的种类最北已分布到北纬34°51′地区,最南已分布北纬23°地区,最东分布到东经120°5′地区。笔者依据的材料仅限于中国分布的喇叭螺属24个物种,其分布亦基本覆盖了此属在中国的分布范围,而且种类数量占该属的80%以上,故可认为基本上反映该属演化和生物地理学变迁的总体格局。

致谢:福建出入境检验检疫局技术中心周卫川研究员协助拍摄标本,特此致谢。
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